Ils sont quatre, héritiers de leurs clans… Ils doivent s’unir pour survivre. La guerre contre les hommes est imminente : les hauts Conseils des Yokaïs se préparent déjà au pire tandis que Bregan, Maya, Nel et Wan associent leurs efforts pour éviter l’inéluctable. Et, alors que le visage de leur véritable ennemi se dévoile, les quatre héritiers n’ont qu’une idée en tête : éviter le bain de sang qui se prépare et protéger ceux qui leur sont chers…

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